Rudolf Harmsen passed away in London, Ont., on March 23, 2022. He was 89. He was predeceased by his brother, Arvid, and is survived by his wife, Jerroldine Harmsen (Arts鈥55); his sister, Miek Zuidema; his sons, Douglas and Leif; son-in-law, Mario Longtin; daughter-in-law, Stephanie Cozart; and granddaughter, Gwendolyn, who knew him as 鈥淕affer.鈥 Rudolf received an undergraduate degree in biology and a master鈥檚 degree in zoology (under zoologist William Beckel) in 1959 from the University of Toronto. He then earned his PhD in zoology in 1963 from Cambridge University, England. For 32 years, from 1966 to 1998, Rudolf was a professor in the Biology Department at 成人大片. He lived an adventure-filled life. His journey as a life-long learner, scientist, educator, husband, father, and grandfather influenced those who came to know him for his loving, generous nature, his profound curiosity, his art, and his wit.